Feng Wen a, ⁎, 1
These authors contribute equally to this work.
, Hai Yu b, e, 1
These authors contribute equally to this work.
, Jinyue Guo a, Yong Li c, Kaijian Luo d, Shujian Huang a, ⁎ 
a College of Life Science and Engineering, Foshan University, Foshan, 528231 Guangdong, China
b Shanghai Veterinary Research Institute, Chinese Academy of Agricultural Sciences, Shanghai 200241, China
c College of Animal Science and Technology, Jiangxi Agricultural University, Nanchang 330045 Jiangxi, China
d College of Veterinary Medicine, South China Agricultural University, Guangzhou 510642 Guangdong, China
e Jiangsu Co-innovation Center for Prevention and Control of Important Animal Infectious Diseases and Zoonoses, Yangzhou 225009, China