The dP4 of Desmanella engesseri Ziegler, 1985Ziegler R. Talpiden (Mammalia Insectivora) aus dem Orleanium und Astaracium Bayerns Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historische Geologie 1985 ;  25 : 131-175

Cliccare qui per andare alla sezione Riferimenti from a maxillary which includes the P2 and P3 in crypt, the P4 in eruption and fully erupted dP4 and M1, is described from Petersbuch 28 (Germany; Lower Miocene, MN3/4). The maxillary was studied using X-ray microtomography to make detailed images of the internal and external features of the sample. This is the first record of a dP4 from Desmanella Engesser, 1972Engesser B. Die Obermiozäne Säugetierfauna von Anwil (Baselland) Tätigkeitsberichte der Naturforschenden Gesellschaft Baselland 1972 ;  28 : 35-363

Cliccare qui per andare alla sezione Riferimenti. Its shape is unique for fossil talpids, possessing the apomorphic feature of replacing the protocone with a large lingual cingulum. The functional use of deciduous teeth and the process that led to the complete loss of milk teeth in extant talpids are discussed.